IHTM15041 | The extent of the share (England, Wales and Northern Ireland): When are the beneficial interests ascertained?
From HM Revenue & Customs · Inheritance Tax Manual
In the case of Pettitt v Pettitt [1970] AC 777 it is considered that
After that those interests remain the same unless and until they are modified by some subsequent agreement either express or to be implied from the circumstances.
You should refer any case where the taxpayer or agent argues that the intention of the parties should be established by examining the whole course of dealings between the parties relevant to the property in question.